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In some cases where a person has lung disease, the partial pressure of oxygen in the pulmonary veins is less than the partial pressure of oxygen in the alveoli.

Which would contribute to the description above?

1) A high proportion of alveoli are collapsed and do not have enough alveolar capillaries.
2) The degree of alveolar ventilation is too low for the cardiac output supplying most of the alveoli.
3) The partial pressure of oxygen in the pulmonary arteries is lower than in the alveolar air.
4) The rate of diffusion of oxygen from the alveolar air to the surrounding alveolar capillaries is too slow.

1 ) 

1, 2 and 4

2 ) 

1, 3 and 4

3 ) 

1 and 2 only

4 ) 

3 and 4 only

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